The SCARABEIDAE Family
Subfamily AEGIALIINAE and APHODIINAE
This preliminary list of Aegialinae and Aphodiinae from Levant was draw up by Giovanni Dellacasa and Marco Dellacasa.
The list is mostly based on bibliographical researches and the reports are referred to more than seven thousand papers on Aphodiinae preserved in author's library. Be aware that it may contain several mistakes of identification, nomenclature and distribution about the taxa involved. For example the checked Agolius bonvouloiri is probably Neocalaphodius moestus, Neagolius praecox is Neagolius ovitensis, Parammoecius corvinus is Parammoecius asphaltinus, etc., etc.
As a matter of fact several taxa need thorough systematic revision considering on "modern" characters, e.g. aedeagus, epipharynx, etc. like for Bodilus nigriventris, Calamosternus pseudounicolor, Phalacronothus hilaris, Chilothorax spp.; Mecynodes spp.; etc. etc.
The nomenclature adopted does not reflect that of "Catalogue of Palaeactic Coleoptera. v. III (2006)" because, at least for Aphodiinae, it results updated to 1999 (first draft supplied). Actually, in the Aphodiinae all the old subgenera of the large genus Aphodius are now upgraded to genus level following "A revisional essay of world genus-group taxa of Aphodiinae..." (Memorie SEI 79, 2001) by Dellacasa G. and alii and "Comments on some systematic and nomenclatural questions..." (Memorie SEI 84, 2005) by M. Dellacasa and G. Dellacasa.
The names of the countries highlighted in bold are those for which the presence of the species has been confirmed on the basis of materials preserved in author's collection.
Subfamily AEGIALIINAE
Tribe EREMAZINI
Genus Eremazus Mulsant, 1851
Eremazus unistriatus Mulsant, 1851 - Israel/Palestine, Jordan, Syria
Subfamily APHODIINAE
Tribe APHODIINI
Genus Acanthobodilus Dellacasa G., 1983
Acanthobodilus immundus (Creutzer, 1799) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Acrossus Mulsant, 1842
Acrossus luridus (Fabricius, 1775) - Israel/Palestine, Syria
Genus Agolius Mulsant and Rey, 1870
Agolius bonvoulori (Harold, 1860) - Syria
Genus Agrilinus Mulsant and Rey, 1870
Agrilinus constans (Duftschmid, 1805) - Lebanon, Syria
Genus Alocoderus A. Schmidt, 1913
Alocoderus hydrochaeris (Fabricius, 1798) - Israel/Palestine, Lebanon, Sinai, Syria
Alocoderus machatschkei (Balthasar, 1960) - Syria
Alocoderus turbatus (Baudi, 1870) - Israel/Palestine, Lebanon, Sinai, Syria
Genus Amidorus Mulsant and Rey, 1870
Amidorus cribrarius (Brulle, 1832) - Israel/Palestine, Jordan, Lebanon, Syria
Amidorus obscurus (Fabricius, 1792) - Israel/Palestine, Syria
Amidorus tarsensis (Petrovitz, 1967) - Lebanon
Amidorus thermicola (Sturm, 1800) - Syria
Genus Ammoecius Mulsant, 1842
Ammoecius naviauxi (Baraud, 1971) - Syria
Genus Aphodius Illiger, 1798
Aphodius fimetarius (Linne, 1758) - Israel/Palestine, Jordan, Lebanon, Syria
Aphodius foetidus (Herbst, 1783) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Biralus Mulsant and Rey, 1870
Biralus equinus (Faldermann, 1835) - Israel/Palestine, Jordan, Lebanon, Syria
Biralus menetriesi (Menetries, 1847) - Israel/Palestine, Jordan, Lebanon, Syria
Biralus satellitius (Herbst, 1789) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Bodiloides G. Dellacasa and M. Dellacasa , 2005
Bodiloides ictericus ssp. ghardimaouensis (Balthasar, 1929) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Bodilopsis Adam, 1994
Bodilopsis rufa (Moll, 1782) - Israel/Palestine
Bodilopsis sordida (Fabricius, 1775) - Israel/Palestine, Syria
Genus Bodilus Mulsant and Rey, 1870
Bodilus longispina (?) (Kuster, 1854) - Israel/Palestine?, Sinai?
Bodilus lugens (Creutzer, 1799) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Bodilus nigriventris (Reitter, 1892) - Syria
Bodilus punctipennis (Erichson, 1848) - Israel/Palestine, Lebanon, Sinai, Syria
Bodilus sabaeus (Peyerhimhoff, 1907) - Israel/Palestine, Jordan, Sinai
Bodilus vartiani (Petrovitz, 1965) - Syria
Genus Calamosternus Motschulsky, 1860
Calamosternus granarius (Linne, 1767) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Calamosternus hyxos (Petrovitz, 1962) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Calamosternus pseudounicolor (Balthasar, 1960) - Israel/Palestine, Syria
Calamosternus trucidatus (Harold, 1863) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Chilothorax Motschulsky, 1860
Chilothorax discedens (A. Schmidt, 1907) - Israel/Palestine, Jordan
Chilothorax distinctus (O.F. Muller, 1776) - Israel/Palestine
Chilothorax equitis (W. Koshantschikov, 1912) - Syria
Chilothorax hieroglyphicus (Klug, 1845) - Israel/Palestine, Jordan, Lebanon, Syria
Chilothorax hucklesbyi (Paulian, 1942) - Israel/Palestine
Chilothorax lineolatus (Illiger, 1803) - Israel/Palestine, Jordan, Lebanon, Syria
Chilothorax melanostictus (W. Schmidt, 1840) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Chilothorax paykulli (Bedel, 1907) - Lebanon, Syria
Chilothorax xanthopterus (Balthasar, 1938) - Lebanon, Syria
Genus Colobopterus Mulsant, 1842
Colobopterus erraticus (Linne, 1758) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Erytus Mulsant and Rey, 1870
Erytus aequalis (A. Schmidt, 1907) - Israel/Palestine, Jordan, Syria
Erytus cognatus (Fairmaire, 1860) - Israel/Palestine, Jordan, Lebanon, Syria
Erytus lindemannae (Balthasar, 1960) - Israel/Palestine, Syria
Erytus mesopotamicus (Petrovitz, 1961) - Israel/Palestine, Syria
Erytus opacior (D. Koshantschikov, 1894) - Israel/Palestine, Jordan
Erytus persicus (Petrovitz, 1961) - Jordan
Erytus pruinosus (Reitter, 1892) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Esymus Mulsant and Rey, 1870
Esymus filitarsis (Reitter, 1898) - Israel/Palestine, Jordan, Lebanon, Syria
Esymus merdarius (Fabricius, 1775) - Israel/Palestine
Esymus ornatulus (Harold, 1866) - Israel/Palestine, Jordan, Lebanon, Syria
Esymus pusillus (Herbst, 1789) - Israel/Palestine
Esymus suturinigra (A. Schmidt, 1916) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Eudolus Mulsant and Rey, 1870
Eudolus quadriguttatus (Herbst, 1783) - Israel/Palestine, Jordan, Syria
Genus Euheptaulacus G. Dellacasa, 1983
Euheptaulacus carinatus (Germar, 1824) - Israel/Palestine, Jordan
Euheptaulacus sus (Herbst, 1783) - Syria
Genus Euorodalus G. Dellacasa, 1983
Euorodalus longevittatus (A. Schmidt, 1916) - Israel/Palestine, Jordan, Lebanon, Syria
Euorodalus tersus (Erichson, 1848) - Syria
Genus Eupleurus Mulsant, 1842
Eupleurus subterraneus (Linne, 1758) - Israel/Palestine?
Genus Grandinaphodius Ziani, 2002
Grandinaphodius inferorum Ziani, 2002 - Lebanon
Grandinaphodius smoliki (Kaufel, 1914) - Israel/Palestine?, Syria
Genus Labarrus Mulsant and Rey, 1870
Labarrus digitatus (Harold, 1871) - Israel/Palestine, Syria
Labarrus lividus (Olivier, 1789) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Labarrus splendidus (Petrovitz, 1955) - Israel/Palestine, Jordan
Genus Liothorax Motschulsky, 1860
Liothorax isikdagensis (Balthasar, 1952) - Israel/Palestine, Syria
Liothorax plagiatus (Linne, 1767) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Loraphodius Reitter, 1892
Loraphodius suarius (Faldermann, 1835) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Loraspis Mulsant and Rey, 1870
Loraspis frater Mulsant and Rey, 1870 - Syria
Genus Mecynodes Mulsant and rey, 1870
Mecynodes angulosus (Harold, 1869) - Israel/Palestine, Jordan, Lebanon, Syria
Mecynodes becvari (Cervenka, 1995) - Jordan
Mecynodes leucopterus (Klug, 1945) - Israel/Palestine, Jordan, Lebanon, Syria
Mecynodes striatulus (Waltl, 1835) - Syria
Mecynodes trochylus (Reitter, 1892) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Megatelus Reitter, 1892
Megatelus contractus (Klug, 1845) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Megatelus scolytiformis (Reitter, 1892) - Israel/Palestine?, Syria?
[syn. Megatelus brahminus Harold, 1879 - Israel/Palestine?, Syria?]
[syn. Megatelus brahminus Harold, 1879 - Israel/Palestine?, Syria?]
Genus Melinopterus Mulsant, 1842
Melinopterus consputus (Creutzer,1799) - Israel/Palestine, Jordan, Lebanon, Syria
Melinopterus prodromus (Brahm,1790) - Israel/Palestine, Jordan, Lebanon, Syria
Melinopterus pubescens (Sturm,1805) - Israel/Palestine, Jordan, Lebanon, Syria
Melinopterus punctatosulcatus ssp. hirtipes (Fischer, 1844) - Israel/Palestine, Jordan, Lebanon, Syria
Melinopterus sphacelatus (Panzer, 1798) - Israel/Palestine
Melinopterus tingens (Reitter, 1892) - Israel/Palestine, Jordan, Lebanon
Genus Mendidaphodius Reitter, 1901
Mendidaphodius angustatus (Klug, 1845) - Israel/Palestine
Mendidaphodius armiger (Harold, 1874) - Israel/Palestine, Syria
Mendidaphodius lepidulus (Harold, 1866) - Israel/Palestine, Jordan, Lebanon
Mendidaphodius linearis (Reiche and Saulcy, 1856) - Israel/Palestine, Jordan, Lebanon, Syria
Mendidaphodius palaestinensis (Petrovitz, 1963) - Israel/Palestine, Jordan
Mendidaphodius rutilus (Klug, 1845) - Sinai, Syria?
Genus Mendidius Harold, 1868
Mendidius assectator (Balthasar, 1961) - Israel/Palestine, Jordan, Syria
Mendidius (?) calliger (Sahlberg, 1908) - Israel/Palestine, Lebanon, Syria
Mendidius fimbriolatus (Mannerheim, 1849) - Israel/Palestine, Jordan, Syria
Mendidius laevicollis (Harold, 1866) - Sinai
Mendidius palmetincolus (Karsch, 1881) - Israel/Palestine, Sinai
Genus Mesontoplatys Motschulsky, 1864
Mesontoplatys arabicus (Harold, 1875) - Israel/Palestine, Sinai
Genus Neagolius W. Koshantschikov, 1912
Neagolius praecox (Erichson, 1848) - Syria?
Genus Nialosternus Hollande and Therond, 1999
Nialosternus sitiphoides (d'Orbigny, 1896) - Israel/Palestine, Jordan, Syria
Genus Nialus Mulsant and Rey, 1870
Nialus politus (Mulsant and Rey, 1870) - Israel/Palestine, Jordan, Lebanon, Syria
Nialus varians (Duftschmid, 1805) - Israel/Palestine, Jordan
Genus Nimbus Mulsant and Rey, 1870
Nimbus contaminatus (Herbst, 1783) - Israel/Palestine
Nimbus harpagonis (Reitter, 1890) - Israel/Palestine, Jordan, Lebanon, Syria
Nimbus libanonensis (Petrovitz, 1958) - Israel/Palestine, Jordan, Lebanon, Syria
Nimbus obliteratus (Panzer, 1823) - Israel/Palestine
Genus Nobiellus M. Dellacasa and G. Dellacasa, 2005
Nobielllus pustulifer (Reitter, 1892) - Lebanon, Syria
Genus Nobius Mulsant and Rey, 1870
Nobius serotinus (Panzer, 1799) - Israel/Palestine
Genus Oxyomus Dejean, 1833
Oxyomus sylvestris (Scopoli, 1763) - Israel/Palestine, Syria
Genus Parabodilus Hollande and Therond, 1999
Parabodilus wollastoni ssp. iranicus (Balthasar, 1946) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Genus Paracoptochirus Balthasar, 1963
Paracoptochirus singularis (Harold, 1868) - Syria
Genus Parammooecius Seidlitz, 1891
Parammoecius corvinus (Erichson, 1848) - Syria
Genus Phalacronothus Motschulsky, 1860
Phalacronothus fumigatulus (Reitter, 1892) - Israel/Palestine/Palestine, Jordan, Lebanon, Syria
Phalacronothus hilaris (Harold, 1869) - Israel/Palestine/Palestine, Jordan, Syria
Phalacronothus quadrimaculatus (Linne, 1761) - Israel/Palestine/Palestine, Jordan, Lebanon, Syria
Genus Pharaphodius Reitter, 1892
Pharaphodius (sensu lato) desertus (Klug, 1845) - Sinai
Genus Plagiogonus Mulsant, 1842
Plagiogonus arenarius (Olivier, 1789) - Syria
Plagiogonus esymoides (Reitter, 1893) - Israel/Palestine
Plagiogonus nanoides (Balthasar, 1961) - Israel/Palestine, Jordan, Syria
Plagiogonus praeustus (Ballion, 1871) - Israel/Palestine, Syria
Plagiogonus syriacus (Harold, 1863) - Lebanon, Israel/Palestine, Syria
Genus Planolinellus M. Dellacasa and G. Dellacasa, 2005
Planolinellus vittatus (Say, 1825) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Pseudacrossus Reitter, 1892
Pseudacrossus koshantschikovi (Jakobson, 1911) - Israel/Palestine, Jordan, Lebanon, Syria
Pseudacrossus suffertus (A. Schmidt, 1916) - Israel/Palestine, Syria
Genus Pseuderytus Hollande and Therond, 1999
Pseuderytus chobauti (Clouet, 1896) - Israel/Palestine, Jordan
Genus Pseudesymus d'Orbigny, 1896
Pseudesymus lucidus (Klug, 1845) - Israel/Palestine, Sinai
Genus Sahlbergianus M. Dellacasa and G. Dellacasa, 2002
Sahlbergianus longissimus (Sahlberg, 1908) - Israel/Palestine, Jordan, Lebanon, Syria
Genus Subrinus Mulsant and Rey, 1870
Subrinus (?) feculentus (Fairmaire, 1892) - Israel/Palestine
Subrinus sturmi (Harold, 1870) - Israel/Palestine
Subrinus vitellinus (Klug, 1845) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Genus Trichonotulus Bedel, 1911
Trichonotulus scrofa (Fabricius, 1787) - Syria
Genus Volinus Mulsant and Rey, 1870
Volinus (?) signifer (Mulsant and Rey, 1870) Israel/Palestine, Syria
Tribe EUPARIINI
Genus Ataenius Harold, 1867
Ataenius horticola Harold, 1869 - Israel/Palestine, Jordan, Lebanon, Syria
Tribe PSAMMODIINI
Genus Brindalus Landin, 1960
Brindalus porcicollis (Illiger, 1803) - Israel/Palestine, Lebanon, Syria
Genus Diastictus Mulsant, 1842
Diastictus vulneratus (Sturm, 1805) - Israel/Palestine, Lebanon
Genus Granulopsammodius Rakovic, 1981
Granulopsammodius plicatulus (Fairmaire, 1892) - Lebanon
Genus Leiopsammodius Rakovic, 1981
Leiopsammodius laevicollis (Klug, 1845) - Israel/Palestine, Jordan, Syria
Genus Psammodius Fallen, 1807
Psammodius asper (Fabricius, 1775) - Israel/Palestine
Psammodius basalis Mulsant and Rey, 1870 - Syria
Psammodius laevipennis A. Costa, 1844 - Israel/Palestine, Lebanon, Syria
Psammodius nocturnus Reitter, 1892 - Israel/Palestine, Lebanon, Syria
Tribe RHYSSEMINI
Genus Pararhyssemus Balthasar, 1956
Pararhyssemus coluber (Mayet, 1887) - Sinai
Genus Platytomus Mulsant, 1842
Platytomus laevistriatus (Perris, 1869) - Israel/Palestine
Platytomus tibialis (Fabricius, 1798) - Israel/Palestine, Lebanon, Syria
Genus Pleurophorus Mulsant, 1842
Pleurophorus anatolicus Petrovitz, 1961 - Israel/Palestine, Jordan, Lebanon, Syria
Pleurophorus arabicus Pittino and Mariani, 1986 - Israel/Palestine, Jordan
Pleurophorus caesus (Creutzer, 1796) - Israel/Palestine, Lebanon, Syria
Pleurophorus opacus Reitter, 1893 - Israel/Palestine, Jordan
Genus Rhyssemodes Reitter, 1892
Rhyssemodes orientalis (Mulsant and Godart, 1875) - Israel/Palestine, Jordan, Lebanon, Sinai, Syria
Genus Rhyssemus Mulsant, 1842
Rhyssemus annaedicatus Pierotti, 1980 - Syria
Rhyssemus beritensis Marseul, 1878 - Israel/Palestine, Jordan, Lebanon, Syria
Rhyssemus geminatus Reitter, 1890 - Lebanon
Rhyssemus germanus (Linne, 1767) - Syria
Rhyssemus granosus (Klug, 1842) - Sinai
Rhyssemus syriacus Petrovitz, 1971 - Jordan, Syria
Rhyssemus verrucosus Mulsant, 1842 - Israel/Palestine
Genus Trichiorhyssemus Clouet, 1901
Trichiorhyssemus setulosus (Reitter, 1892) - Israel/Palestine
Selected References
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